But centre Gavin Henson has not been included among coach Warren Gatland's matchday squad for the game at Twickenham.
Scarlets fly-half Jones makes his 101st Test appearance, putting him one in front of Gareth Thomas.
Elsewhere, Gatland hands starts to full-back Rhys Priestland, 19-year-old George North on the right wing and Huw Bennett at hooker instead of an injured Matthew Rees.
Flanker Sam Warburton captains the team in Rees' absence, but Henson will have to wait to press any World Cup selection claims. Wales meet England in Cardiff on Saturday week, then host Argentina seven days later.
Gatland plans to name his final 30-man squad that will travel to New Zealand on August 22.
Henson is replaced in the side by Jamie Roberts, with Wales coach Warren Gatland admitting he has opted for his strongest available line-up.
"These are the men in possession of the jersey at the moment and it is now up to them to hold on to it and they will only do that with a performance at Twickenham at the weekend," he said.
"The stakes are high for individuals, but as a team it is also important for us to create some momentum as we head for the World Cup in New Zealand.
"We have made changes but we have some pretty established combinations coming back with Jamie Roberts returning to the midfield and Alun-Wyn (Jones) and Bradley (Davies) in the second row.
Assistant coach Rob Howley praised Stephen Jones' record-breaking achievement, although he playfully suggested the focus of attention should fall on Alun Wyn Jones' half-century of caps, which he will bring up in London.
"It is a tremendous achievement for Stephen, but he had all the well deserved accolades when he won his 100th cap and the spotlight this weekend should really go to Alun-Wyn on winning his 50th," he said.
Hawley also backed full-back Priestland to impress with the backline experience of Roberts and Shane Williams alongside him.
"We are fielding nine of the players who played in the final 6 Nations match this year and have some senior men coming back in with Jamie and Shane bringing a wealth of experience," he added.
"At full-back Rhys has done very well for the Scarlets and we have an opportunity to test him now at this level and we are sure he won't let anyone down."
England: D Armitage (London Irish); M Banahan (Bath Rugby), M Tuilagi (Leicester Tigers), R Flutey (London Wasps), M Cueto (Sale Sharks); J Wilkinson (Toulon), D Care (Harlequins); A Corbisiero (London Irish), D Hartley (Northampton Saints), M Stevens (Saracens), S Shaw (Unattached), T Palmer (Stade Francais), T Croft (Leicester Tigers), L Moody (Bath Rugby, capt), J Haskell (Ricoh Black Rams).
Replacements: L Mears (Bath Rugby), D Wilson (Bath Rugby), M Botha (Saracens), T Wood (Northampton Saints), R Wigglesworth (Saracens), C Hodgson (Saracens), C Sharples (Gloucester Rugby).
Wales: R Priestland (Scarlets); G North (Scarlets), J Davies (Scarlets), J Roberts (Cardiff Blues), S Williams (Ospreys); S Jones (Scarlets), M Phillips (Bayonne); P James (Ospreys), H Bennett (Ospreys), C Mitchell (Exeter), B Davies (Cardiff Blues), A-W Jones (Ospreys), D Lydiate (Newport Gwent Dragons), S Warburton (Cardiff Blues, capt), T Faletau (Newport Gwent Dragons).
Replacements: L Burns (Newport Gwent Dragons), R Bevington (Ospreys), L Charteris (Newport Gwent Dragons), R Jones (Ospreys), T Knoyle (Scarlets), S Williams (Scarlets), M Stoddart (Scarlets).
Gilbert has released a new line of rugby cleats. The Gilbert Virtuo 8S is part of the exciting new product. Check it out.
The entire All Blacks apparel line has been updated for 2013/14. Check out the New Zealand All Blacks polo.
The Nike Tiempo is a solid rugby cleat and one of few styles still made from full-grain natural leather.
The Gilbert Blitz 8S rugby cleat is a great cleat at a great price of $69.99. Get a new pair of cleats today.
A cool looking all black rugby cleat with the high performance adidas is known for. Get in the Gear!
Wear the crest of the British and Irish Lions on your t-shirt. A great look for the summer.
The All Blacks Performance t-shirt is black with hints of blue from the training jersey. Very Cool.
The New Zealand All Blacks training jersey for 2013/14. Get in the Gear!
The USA Rugby Pro Alternate rugby jersey is perfect for any fan of the Eagles. Get yours to wear during the summer Test matches.
The NEW All Blacks 2013/14 jersey has arrived at World Rugby Shop. Dare to wear the colors of the All Blacks.